Graduate Course Listings

Note: Subject to change. Please refer to the Graduate Calendar for confirmation of fall/winter offerings listed below.

Biol-860-Biol-865 inclusive, are modules worth 1.5 credit units each.  These courses are also CAPPED at 12 and require written permission from the course coordinator.  Only the Biology Graduate  Assistant will register you into any of these courses once the permission has been received by her.   

All other courses are half-courses (3.0 credit units) which are offered either in the fall or winter term.  


QACS 799
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QACS 799 Introduction to Animal Care - WebCT

Anyone who is going to be listed on an Animal Use Protocol (AUP) has to take the course. This pertains to Principal Investigators as well as any individuals who are responsible for the care and use of animals under an approved AUP. 

Link:  :

Fall / Winter / Spring Animal Care Service
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Biol-812 Programming for Biologists

Description/Outline:  Programming for Biologists

This hands-on course introduces students to modern computational methods for biology with an emphasis on emerging standards of Open Science and Reproducible Research. No prior programming experience is assumed. In coordination with Queen’s University Centre for Advanced Computing (CAC), the content of this course is intended to teach core-competency in the following areas:

1. Programming basics: regular expressions, flow control, file manipulation, custom functions.

2. Unix/Linux command-line programming and shell scripting on CAC infrastructure (SLURM).


R with GitHub, R Markdown, and the tidyverse (dplyr, %>%, tidyr, ggplot2) for seamless reproducible data management, analysis and production of publication-ready graphics.

3. Introduction to some standard formats (e.g. FASTA, FASTQ, SAM, BED, BAM), programs (e.g. SAMTOOLS, VCF, HISAT2), and pipelines in Unix/Linux, R and Python for reproducible analysis of data from next-generation sequencing platforms.


Content includes introduction to state-of-the-art methods in genomics and metagenomics, including sequence assembly, alignment, variant detection, and gene annotation. However, the majority (~3/4) of the course is devoted to learning basic computational tools that may be useful to biologists working on large datasets of any kind.

Winter 2018 R. Colautti
BIOL 819
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Biol 819 Selected Topics in Molecular Biology

Topics vary annually to reflect the latest status of the field of molecular genetics .

Winter 2018 W. Bendena
BIOL 820
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Biol 820 Commercialization of Biological Research
Current issues relating to the biotechnology industry will be dealt with in detail. Topics covered include: grant writing; patenting; circumventing patents; funding sources; business plans; venture capital investments; public awareness; public perspective; and layperson presentations. Three term hours; fall. K. Ko and V.K. Walker. PREREQUISITE: At least one of the following: BIOL-201*, BIOL-205*, MBIO-318*, BIOL-441*, BIOL-330*/430* or equivalent.
Fall 2017 K. Ko
BIOL 822
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Biol-822 Long-Term Environmental Change

The main focus of this course will be to review and assess the many techniques currently available to track long-term environmental change. An emphasis will be placed on biological approaches dealing with sedimentary analyses, but other proxy methods (e.g. ice cores, bore holes, etc.) will also be covered. General topics to be covered will include climatic change, acidification, eutrophication, lake and reservoir management, UV penetration, etc. Three term hours.  Fall.  J.P. Smol 

Fall 2017 John Smol
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Biol-824 Gateway to Graduate Studies in Biology
No description available.
Fall 2017 F. Bonier
BIOL 848
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Biol 848 Field Courses in Biology
No description available.
Fall Not Offered
Winter Lougheed
Spring Wang
BIOL 849
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Biol 849 Environmental Issues

Consideration will be given to environmental, legal, economic, political, sociological and biological aspects of current issues in the management of the Great Lakes. Models for managing nutrients, toxics and fisheries will be compared from a multidisciplinary viewpoint. Three term hours; Fall.  B. Zeeb

Fall 2017 Barb Zeeb
BIOL 860
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Biol 860 Introduction to Statistics


This course is for new graduate students who are at early stages of planning their research projects and collecting data. Topics will include experimental design, matching hypotheses with appropriate statistical analyses, parameter estimation, and graphing. Statistical analyses will be based on a foundation using normal error distribution. Students will be introduced to R, a Language and Environment for Statistical Computing. 1.5 credit units.

Fall 2017 S. Arnott
BIOL 861
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Biol 861 Introduction to Linear Models

This course is designed for Biology graduate students with a basic introductory statistics/experimental design course and a working knowledge of R, a Language and Environment for Statistical Computing. In-depth exploration of all aspects of fitting linear models to continuous and categorical data, using mainly the lm function in R. Topics include residual analysis, maximum-likelihood methods, graphical presentations, ordinary least squares, model II regression, transformations, model selection with focus on information-theoretic approaches and outlier detection. 1.5 credit units.

Lecturer: C. Eckert

Fall 2017 C. Eckert
BIOL 862
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Biol 862 Generalized Linear Models

This is a course designed to introduce students the linear model for analyzing continuous and categorical data in biology. Each session will begin with a short lecture by the faculty instructor, followed by a student presentation of homework, and a hands-on interactive session focusing on statistical computing and graphics. Each student will bring a laptop to class.

Winter 2018 R. Montgomerie
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Biol-893 Mentoring Experience in Biology


Students will advise and train other students in biological investigations, normally over a two-term period.



Activities include guidance on research proposals, research procedures, student presentations, and drafts of student work



Assessment of mentoring skills as you provide guidance on research proposals, research procedures, student presentations, and drafts of student work






Open to full-time students having completed two terms of study in Biology M.Sc. or Ph.D. programs.





Does not count towards degree

Fall 2017 and Winter 2018 Your Supervisor
BIOL 897
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Biol 897 Seminars in Biology

Attending a diverse array of seminars is an essential component in the development of a student.  The aim of this course is to develop skills in listening, synthesizing and critical thinking, as well as fostering the development of important oral and written communication skills.  Students are required to attend and evaluate departmental or specialized research seminars as well as to present a seminar based on their graduate thesis research.  Fall/Winter/Summer. L. Seroude & P. Martin.

Information and instructions for graduate students of MCIB faculty:

Questions to Dr. Laurent Seroude (

Information and instructions for graduate students of EEB faculty:

Questions to Dr. Paul Martin (

Enrolment is extended over a period of SIX terms ONLY.  

Students must register in two academic years.  Ensure that you email the Biology Graduate Assistant (Joanne Surette) to register you for the second year.



Fall / Winter / Summer L. Seroude P. Martin
Biol-951 and Biol-952
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Biol-951 Biol-952 Advanced Studies in Ecology, Evolution and Behaviour (50 Years of Invasion Genetics)

Invasion genetics is the study of genetic diversity and evolutionary processes involving invasive species. It covers a range of topics from the biological characteristics of successful invaders to the evolutionary consequences of biological invasions. This course examines current topics in invasion genetics and their historical origins. Students will present and discuss papers from the 2015 book “Invasion Genetics: The Baker & Stebbins Legacy” and contrast these with related chapters from the 1965 book “The Genetics of Colonizing Species.”  A concise final paper will review the history and ‘state of the art’ of a main questions in invasion genetics, and identify open questions that have not received as much attention.

cancelled cancelled
BIOL 953 & 954
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Biol 953 & 954 Advanced Studies in Plant Sciences

An investigation of recent topics in plant molecular biology and biochemistry.  Topics will selected according to student interest but are likely to include adaptation to stress, protein structure/function, development, and plant biotechnology. Material will be drawn from recent primary literature.

Winter 2018 W. Snedden
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BIOL 955 Cell Cycle

The cell cycle, its major periodic events, the G0-G1 transition, and the integration of growth and cell division will be the major foci.  The course will consider the historical origins of the field as well as the modern integration of genetics, cell and molecular biology.   Material will be drawn mostly from the research literature last few years.  

Winter 2018 P. Young
BIOL 956
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Biol 956 Advanced Studies in Molecular & Genetics

This is the introductory course for graduate students in the Cellular, Molecular and Integrative Biology (CMIB) program.  This course provides graduate students with a solid function of molecular and cell biology. Rather than offer small, specialized courses with very few students, the intent is to cover a broad field and to provide an opportunity for students with varying interests to interact.  Peer interaction is necessary in an intellectual environment. The course will appeal to most disciplines and not be constrained by a single model system, biological process, or level of organization (genomic, cell, biochemistry, etc). Subjects will be chosen based on students' interests.

Winter 2018 I. Chin-Sang

I've spent more time than many will believe [making microscopic observations], but I've done them with joy, and I've taken no notice those who have said why take so much trouble and what good is it?

Antonie van Leeuwenhoek

It's a parts list... If I gave you the parts list for the Boeing 777 and it had 100,000 parts, I don't think you could screw it together and you certainly wouldn't understand why it flew

Eric Lander

What is true for E. coli is also true for the elephant

Jacques Monod

The world becomes full of organisms that have what it takes to become ancestors. That, in a sentence, is Darwinism

Richard Dawkins

Shall we conjecture that one and the same kind of living filaments is and has been the cause of all organic life?

Erasmus Darwin

Nature proceeds little by little from things lifeless to animal life in such a way that it's impossible to determine the line of demarcation


Cells let us walk, talk, think, make love, and realize the bath water is cold

Lorraine Lee Cudmore

In the distant future I see open fields for far more important researches. Psychology will be based on a new foundation, that of the necessary acquirement of each mental power and capacity by gradation. Light will be thrown on the origin of man and his history

Charles Darwin

It is my belief that the basic knowledge that we're providing to the world will have a profound impact on the human condition and the treatments for disease and our view of our place on the biological continuum

J. Craig Venter

Imagine a house coming together spontaneously from all the information contained in the bricks: that is how animal bodies are made

Neil Shubin

A grain in the balance will determine which individual shall live and which shall die - which variety or species shall increase in number, and which shall decrease, or finally become extinct

Charles Darwin

The stuff of life turned out to be not a quivering, glowing, wondrous gel but a contraption of tiny jigs, springs, hinges, rods, sheets, magnets, zippers, and trapdoors, assembled by a data tape whose information is copied, downloaded and scanned

Steven Pinker

We wish to discuss a structure for the salt of deoxyribose nucleic acid. (D.N.A.). This structure has novel features which are of considerable biologic interest

Rosalind Franklin

We are biology. We are reminded of this at the beginning and the end, at birth and at death. In between we do what we can to forget

Mary Roach

The systems approach to biology will be the dominant theme in medicine

Leroy Hood

I've always been interested in animal behavior, and I keep reading about it because it's so surprising all the time - so many things are happening around us that we neglect to look at. Part of the passion I have for biology is based on this wonderment"

Isabella Rossellini

Because all of biology is connected, one can often make a breakthrough with an organism that exaggerates a particular phenomenon, and later explore the generality

Thomas Cech

Nothing in Biology makes sense except in the light of evolution

Theodosius Dobzhansky

Biology is now bigger than physics, as measured by the size of budgets, by the size of the workforce, or by the output of major discoveries; and biology is likely to remain the biggest part of science through the twenty-first century

- Freeman Dyson

Nothing can be more incorrect than the assumption one sometimes meets with, that physics has one method, chemistry another, and biology a third

- Thomas Huxley